of the Subclass Prototheria (all other living mammals belong to the subclass Theria). McKenna and Bell (1979) divided the order into two (Platypoda and Tachyglossa); the date of divergence of the two living families is unknown, and conservatively they are retained here in a single order.
Reading on, I realise weâre talking about echidnas and platypuses, but I realise also, from the profound depths of my ignorance, that scientific detail is going to be hard on the digestion. Then â glory! â I remember Alfred Russel Wallace, for whom much of the world truly was a blank canvas, and I return to him as to a kind of intellectual comfort blanket. Let me begin where he began, and be led from there through zoological history.What were the âvery peculiar forms of mammaliaâ that struck him so forcefully in tropical and southern Africa? âSuch are the golden moles, the Potamogale, and the elephant-shrews . . .â
Golden moles . I turn back to Mammal Species of the World , and claw my way to page 77:
SUBORDER CHRYSOCHLORIDEA Broom, 1915
COMMENTS: MacPhee and Novacek (1993) erected the suborder Chrysochloromorpha for golden moles, but following Simpsonâs (1945: 32â33) nomenclatural principles for categories above superfamilies, Chrysochloridea is the senior synonym.
Chrysochloridea it is, then. There are a good few of them â many more, I suspect, than even Wallace would have imagined. Most, but not all, have common as well as Latin names, usually in honour of their discoverers, territories or physical peculiarities. First up is Arendâs golden mole, then Duthieâs, Sclaterâs, Cape, Stuhlmannâs, Visagieâs, Giant, Rough-haired, De Wintonâs, Van Zylâs, Grantâs, Fynbos, Hottentot, Marleyâs, Robust, Highveld, Congo, Yellow, Somali, Gunningâs and Julianaâs. Their territories range all the way down from the Gulf of Guinea, scene of Hannoâs first brush with the Gorillae , through equatorial and sub-equatorial Africa to the Cape. But there are two exceptions, which, weirdly, appear to have no ranges at all.
Visagieâs golden mole ( Chrysochloris visagiei ) â âknown only from the holotypeâ.
Somali golden mole ( Calcochloris tytonis ) â âknown only from the type specimenâ.
By now I know that âholotypeâ and âtype specimenâ are the same thing. In each case they mean the original collected example from which the species was first described and introduced toscience. What we are being told is that, throughout the whole of the scientific age, Visagieâs and the Somali golden moles have each been seen only once. One animal constitutes the entire species. Conservatively, their status is recorded as âcritically endangeredâ. I will discover later that, though this degree of rarity is not a common phenomenon, it is not a rare one either. An astonishing number of species are accorded their identity on astonishingly sparse scraps of evidence. I turn next to the world authority on extinction and survival, the IUCN (International Union for Conservation of Nature) Red List of Threatened Species .
It confirms that Visagieâs golden mole is known from a single specimen collected from the Northern Cape and described in 1950. We do not learn whether the animal was alive or dead, or even complete in all its parts, but a few drops of scepticism leak through the authorâs dry academic prose. âSeveral field trips to ground-truth the occurrence of this species have yielded no specimens, or even signs of golden moles, suggesting either an error in recording provenance, or that the original specimen was transported there by anthropogenic means or even perhaps floodwaters of the Renoster River . . .â
If that is peculiar enough, then itâs nothing to compare with its Somali cousin. Again the Red List confirms the uniqueness of the specimen, found at Giohar, Somalia, in 1964. But